CLADOENDESIS OF EPHEMEROPTERA
Tetramerotarsata Liberevenata Turbanoculata
pm.Protopatellata - Demoulinia)
Nomen hierarchicum: Demoulinia/g(1)
In circumscription fits:
gen. Demoulinia Gillies 1990:97-128: 125.
References. Crass 1947 (Centroptilum pulchrum): ; Gillies 1990:97-128: ;
Autapomorphies of Demoulinia.
(1) In larva, patella-tibial suture is lost on all legs. Belonging of this taxon to Protopatallata [whose larva and female initially lack patella-tibial suture on fore legs and have this suture on middle and hind legs see Protopatellata] is proven by leg structure of winged females: in female subimago and imago partella-tibial suture is clearly developed on middle and hind legs and is entirely absent on fore legs (as in winged male).
(2) Labial palp (retaining all 3 segments and muscle moving 3rd segment) has peculiar shape: its 2nd segment bears large apicomedial projection of nearly quadrate form; more distal angle of this projection is rounded, and more proximal angle bears small point (Wuillot & Gillies 1993: Fig.21).
() Last segment of gonostylus has unusual hooked shape, with wide base and pointed apex (Crass 1947:fIG. 27g).
Characters of Demoulinia of unclear phylogenetic status.
() Mandibles of «Centroptilum-type»: kinetodontium is separated from incisor (but immovable) and rotated perpendicular to the plane of mandible, so that its denticles hide one another in dorsal or ventral view; left prostheca is bifurcate and rotated in the same manner (Fig. ) (see Index of characters [1.1.22], [1.1.24]).
() Hind wings are lost (among Protopatellata the same in Indocloeon, part of Rhithrocloeon/fg1 and part of Anafroptilum see Index of characters [2.2.59]).
() Tergalii are able to make rhythmical respiratory vibration (see Index of characters [1.3.30]).
() Gonovectes are fused with penial bridge. Among Protopatellata, the same in Anafroptilum and Rhithrocloeon/fg1 see Index of characters [2.3.16a].
Plesiomorphies of Demoulinia.
Size. Fore wing length 68 mm (see Tetramerotarsata).
Distribution. South Africa and Madagascar.
Nominal species in Demoulinia/g(1):
assimilis Gattolliat 2003 [Demoulinia]
crassi Demoulin 1971 [Centroptilum] typus nominis Demoulinia --/
insularis Lugo-Ortiz & McCafferty 1998 [Demoulinia]
Turbanoculata INCERTAE SEDIS