CLADOENDESIS OF EPHEMEROPTERA

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Demoulinia

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata Liberevenata Turbanoculata 
 pm.Protopatellata - Demoulinia)

Nomen hierarchicum: Demoulinia/g(1)

In circumscription fits:

— gen. Demoulinia Gillies 1990:97-128: 125.


References. Crass 1947 (Centroptilum pulchrum): ; – Gillies 1990:97-128: ;


Autapomorphies of Demoulinia.

(1) In larva, patella-tibial suture is lost on all legs. Belonging of this taxon to Protopatallata [whose larva and female initially lack patella-tibial suture on fore legs and have this suture on middle and hind legs – see Protopatellata] is proven by leg structure of winged females: in female subimago and imago partella-tibial suture is clearly developed on middle and hind legs and is entirely absent on fore legs (as in winged male).

(2) Labial palp (retaining all 3 segments and muscle moving 3rd segment) has peculiar shape: its 2nd segment bears large apicomedial projection of nearly quadrate form; more distal angle of this projection is rounded, and more proximal angle bears small point (Wuillot & Gillies 1993: Fig.21).

() Last segment of gonostylus has unusual hooked shape, with wide base and pointed apex (Crass  1947:fIG. 27g).

Characters of Demoulinia of unclear phylogenetic status. 

() Mandibles of «Centroptilum-type»: kinetodontium is separated from incisor (but immovable) and rotated perpendicular to the plane of mandible, so that its denticles hide one another in dorsal or ventral view; left prostheca is bifurcate and rotated in the same manner (Fig. ) (see Index of characters [1.1.22], [1.1.24]).

() Hind wings are lost (among Protopatellata the same in Indocloeon, part of Rhithrocloeon/fg1 and part of Anafroptilum – see Index of characters [2.2.59]).

() Tergalii are able to make rhythmical respiratory vibration (see Index of characters [1.3.30]).

() Gonovectes are fused with penial bridge. Among Protopatellata, the same in Anafroptilum and Rhithrocloeon/fg1 – see Index of characters [2.3.16a].

()

Plesiomorphies of Demoulinia.

()

Size. Fore wing length 6–8 mm (see Tetramerotarsata).

Distribution. South Africa and Madagascar.


Nominal species in Demoulinia/g(1):

assimilis Gattolliat 2003 [Demoulinia]

crassi Demoulin 1971 [Centroptilum] — typus nominis Demoulinia --/

insularis Lugo-Ortiz & McCafferty 1998 [Demoulinia]


Examined also:

 


See also:

Turbanoculata INCERTAE SEDIS