CLADOENDESIS OF EPHEMEROPTERA

CLADOENDESIS OF EPHEMEROPTERA
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Callibaetis/fg3

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta
Tetramerotarsata Liberevenata Turbanoculata
Anteropatellata Baetovectata Callibaetis/fg1 Callibaetis/fg2 - Callibaetis/fg3)

Nomen hierarchicum: Callibaetis/fg3 [f:1973; g:1881] (sine Callibaetoides)

In circumscription fits:

— gen. Callibaetis Eaton 1881: 196

— subfam. Callibaetinae: Gillies 1991: 181

Nominal taxa included:

Abaetetuba/g []

Aiso/g []

Cunhaporanga/g []

Neobaetis/g [g:1924]

References. Needham & Traver & Hsu 1935: * *; – Edmunds & Jensen & Berner 1976: * *; – Gillies 1991:VI.Conf.Eph.: * *; – Hofmann & Sartori & Thomas 1999: * *; – Dominguez & Molineri & Pescador & Hubbard & Nieto 2006: * *; – Nieto 2008:229-243: * *

Autapomorphies of Callibaetis/fg3.

(1) In imago of both sexes, thorax and abdomen with irregularly situated dark dots, in the middle of each dot a small seta in light pore. In other mayflies such dots and setae are absent or invisible.

(2) Imaginal fore wing often with dark colour pattern more developed in female than in male (often in male being completely absent): anterior part of wing is coloured, with colourless spots around cross veins. The same in many Cloeon/fg2.

(3) Maxillary palp is 2-segmented. Non-unique apomorphy (see Index of characters [1.1.42]).

(4) Larval abdominal terga lost wide scales [see Turbanoculata (12)] (see Index of characters [1.3.4]).

Size. Fore wing length 6–12 mm (see Tetramerotarsata).

Distribution. America (Nearctic and Neotropical Regions).

Nominal species in Callibaetis/fg3:

abundans Navás 1912 [Baetis] nomen dubium

alegre Traver 1944 [Callibaetis] — syn.subj. willineri [Callibaetis]

americanus Banks 1900 [Callibaetis] — syn.subj. hageni [Callibaetis]

amoenus Navás 1930 [Callibaetis] — syn.subj. zonalis [Callibaetis]

apertus Navás 1917 [Callibaetis] — syn.subj. zonalis [Callibaetis]

apicatus Navás 1917 [Callibaetis] — syn.subj. guttatus [Callibaetis]

brevicostatus Daggy 1945 [Callibaetis] — syn.subj. skokianus [Callibaetis]

bruchius Navás 1920 [Callibaetis] — syn.subj. guttatus [Callibaetis]

californicus Banks 1900 [Callibaetis]

calophenigyn Cruz & Salles & Hamada 2017 [Callibaetis (Aiso)]

calloventer Cruz & Salles & Hamada 2014 [Callibaetis] — typus nominis Cunhaporanga

camposi Navás 1930 [Callibaetis]

capixaba Cruz & Salles & Hamada 2009 [Callibaetis] --

carolus Traver (in Needham & Traver & Hsu) 1935 [Callibaetis] — syn.subj. hageni [Callibaetis]

centralis Peters 1959 [Callibaetis] — syn.subj. pictus Eaton 1871 [Baetis]

claudiae Edmunds 1945 [Callibaetis] nomen nudum — syn.subj. hageni [Callibaetis]

coloradensis Banks 1900 [Callibaetis] — syn.subj. hageni [Callibaetis]

completa Banks 1930 [Callibaetis] — syn.subj. floridanus [Callibaetis]

cruentus Cruz & Salles & Hamada 2014 [Callibaetis] --

depressus Navás 1922 [Callibaetis] — syn.subj. zonalis [Callibaetis]

distinctus Lugo-Ortiz & McCafferty 1996 [Callibaetis]

doddsi Traver (in Needham & Traver & Hsu) 1935 [Callibaetis] — syn.subj. pictus [Baetis]

dominguezi Gillies1990 [Callibaetis]

evergreenensis Thew 1959 [Callibaetis] — syn.subj. hageni [Callibaetis]

fasciata Pictet 1843 [Cloe] — typus nominis Abaetetuba

ferruginea Walsh 1862 [Cloe] --

floridanus Banks 1900 [Callibaetis] --

fluctuans Walsh 1862 [Cloe]

fluminensis Cruz & Salles & Hamada 2009 [Callibaetis] — syn.subj. zonalis [Callibaetis]

fusca Dodds 1923 [Callibaetis] — syn.subj. hageni [Callibaetis]

gelidus Cruz & Salles & Hamada 2014 [Callibaetis]

gloriosus Navás 1923 [Baetis] — syn.subj. sellacki [Cloe]

gonzalezi Navás 1934 [Cloeon] --

gregarius Navás 1930 [Callibaetis]

grisoli Navas [Cloeon] nomen nudum

guttatus Navás 1915 [Callibaetis]

hageni Eaton 1885 [Callibaetis] --

hebes Upholt 1936 [Callibaetis] — syn.subj. hageni [Callibaetis]

imperator Cruz & Salles & Hamada 2017 [Callibaetis (Cunhaporanga)]

itannae Cruz & Salles & Hamada 2014 [Callibaetis] — typus nominis Aiso

jaffueli Navás 1918 [Callibaetis]

jocosa Navás 1912 [Callibaetis]

lineatus Navás 1933 [Callibaetis] — syn.subj. sellacki [Cloe]

lorentzii Weyenbergh 1883 [Cloe] — syn.subj. sellacki [Cloe]

molinerii Cruz & Salles & Hamada 2017 [Callibaetis (Callibaetis)]

montanus Eaton 1885 [Callibaetis]

nigracyclus Cruz & Salles & Hamada 2014 [Callibaetis]

nigrita Banks 1918 [Callibaetis] — syn.subj. hageni [Callibaetis]

nigrivenosa Banks 1918 [Callibaetis]

opacus Navás 1915 [Baetis] — syn.subj. zonalis [Callibaetis]

pacifica Seemann 1927 [Callibaetis] — syn.subj. pictus Eaton 1871 [Baetis]

pallidus Banks 1900 [Callibaetis]

paulinus Navas 1924 [Neobaetis] — typus nominis Neobaetis

pictus Eaton 1871 [Baetis] — typus nominis Callibaetis

pollens Needham & Murphy 1924 [Callibaetis]

pretiosus Banks 1914 [Callibaetis]

punctilusus McCafferty & Provonsha 1993 [Callibaetis montanus]

radiatus Navás 1920 [Callibaetis]

rimatus Navás 1933 [Callibaetis] — syn.subj. jocosa [Callibaetis]

sellacki Weyenbergh 1883 [Cloe]

semicostata Banks 1914 [Callibaetis] — syn.subj. pallidus [Callibaetis]

signata Banks 1918 [Callibaetis] — syn.subj. pictus [Baetis]

skokiana Needham 1903 [Callibaetis]

sobrius Navás 1916 [Callibaetis] — syn.subj. zonalis [Callibaetis]

spegazzinus Navás 1920 [Callibaetis] — syn.subj. jocosa [Callibaetis]

stictogaster Navás 1915 [Callibaetis]

tessellata Hagen 1861 [Baetis] nom.praeocc. — syn.obj. hageni [Callibaetis]

traverae Upholt 1937 [Callibaetis] — syn.subj. fluctuans [Cloe]

trifasciatus Esben-Petersen 1912 [Callibaetis] — syn.subj. fasciata [Cloe]

undata Pictet 1843 [Cloe]

venulosus Navás 1933 [Callibaetis] — syn.subj. radiatus [Callibaetis]

viparius Needham & Murphy 1924 [Callibaetis]

virellus Navás 1915 [Baetis] — syn.subj. zonalis [Callibaetis]

vitrea Dodds 1923 [Callibaetis] nom.praeocc. — syn.obj. doddsi [Callibaetis]

vitreus Navás 1915 [Callibaetis]

vitreus Navás 1919 [Callibaetis] nom.praeocc. — syn.subj. zonalis [Callibaetis]

viviparus Needham & Murphy 1924 [Callibaetis]

willineri Navás 1933 [Callibaetis]

zonalis Navás 1915 [Callibaetis]

zonatus Navás 1929 [Callibaetis] — syn.subj. guttatus [Callibaetis]

Examined also:

sp.1 (Peru) --

sp.2 (Peru)

sp.3 (Peru) /

sp.4 (Peru)

sp.n.5 (Peru) --

sp.n.6 (Peru) --