CLADOENDESIS OF EPHEMEROPTERA

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Callibaetis/fg1

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata Liberevenata Turbanoculata 
Anteropatellata Baetovectata - Callibaetis/fg1)

Nomen hierarchicum: Callibaetis/fg1 [f:1973; g:1881] (incl. ...., Callibaetoides)

In circumscription fits:

 


References.  


Autapomorphies of Callibaetis/fg1.

(1) Labrum and hypopharynx form a peculiar grasping apparatus: Labrum has a pair of sclerotized protuberances on sides of its median incision [see Turbanoculata (4)]; apex of hypopharynx has an unpaired sclerotized protuberance on dorsal side; when labrum and hypopharynx are pressed together, the sclerotized protuberance of hypopharynx is inserted between the sclerotized protuberances of labrum. Usually the protuberances of labrum are distinctly outlined, but in floridanus [Callibaetis] they are fused with other sclerotization of distal margin of labrum.

(2) Mandibles characteristic and uniform in all representatives: outer margin laterad of incisor convex, kinetodontium stout but well-separated, left prostheca stout and generally bifid (sometimes with smaller apical denticles), right prostheca slender, bifid and straight (not markedly bent proximally).

(3) Maxilla always has characteristic shape: relatively short, with especially convex outer margin.

(4) Labial palp has boundary between 2nd and 3rd segments oblique, so that 2nd segment is long at outer side and short at inner side; 2nd and 3rd segments together form integral figure. 

Characters of Callibaetis/fg1 of unclear phylogenetic status. 

(5) Unlike other Turbanoculata, in Callibaetis/fg1 external larval gonostyli buds are long, projected far behind sternum IX; this resembles a plesiomorphic condition present in Siphlaenigma and majority of other Ephemeroptera. In mature larva ready to moult to subimago, buds of subimaginal gonostyli are folded under larval cuticle ?similar to the "Nigrobaetis-type":??? 2nd segment of gonostylus is directed posteriorly and curved by its convexity medially and by apex laterally [see Baetovectata (3)] (unlike Baetofemorata and Labiobaetis/f1 see Index of characters [2.3.9]).

(6) Larval antennae bases are not brought together, frons between them does not form a keel (unlike Paracloeodes and some Baetungulata see Index of characters [1.1.3]).

(7) Tergalii are able to make rhythmical respiratory movements (see Index of characters [1.3.30]).

Plesiomorphies of Callibaetis/fg1.

Hind wings are always developed [being strongly diminished see Turbanoculata (10)]. 

Larva always of siphlonuroid (actively-swimming) type: legs slender, not tenacious; abdomen large, not flattened ventrally (terga not wider than sterna), paracercus and primary swimming setae well-developed. Larval claw with 2 rows of denticles [see Tetramerotarsata (11); see bellow, Callibaetis/fg2 (3)].

Variable characters of Callibaetis/fg1. On fore wing intercalaries are situated mainly in twos in each field [see Baetovectata (1)]; sometimes one of two veins is lost, and in each field a single intercalary is present, being situated either eccentrically, or in centre of field; sometimes male has two intercalaries, and female has one intercalary in each space.

Size. Fore wing length 412 mm (see Tetramerotarsata).

Distribution. America (Nearctic and Neotropical Regions).


The taxon Callibaetis/fg1 is divided into:

1. ... 

       collendum Harker 1957 [Centroptilum]

       elongatum Suter 1986 [Centroptilum]

2. Callibaets/fg2

2.1. Callibaetoides 

2.2. Callibaets/fg3