CLADOENDESIS OF EPHEMEROPTERA

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Baetiella/g1

(Panephemeroptera Euephemeroptera Euplectoptera Anteritorna pm.Tridentiseta  
Tetramerotarsata Liberevenata Turbanoculata  
Anteropatellata Baetovectata Baetungulata Baetofemorata - Baetiella/g1)

Nomen hierarchicum: Baetiella/g1 [g:1931] (incl. Neobaetiella)

In circumscription fits:

— subgen. Baetiella: Kazlauskas 1963: 586

— gen. Baetiella Ueno 1931: 220

Synonyms: subjective synonym of  Baetiella/g [g:1931] (typus Acentrella japonica) is Matsumuracloeon/g [g:1989] (typus Pseudocloeon aino) (nom. pro Pseudocloeon Matsumura 1931: 1473, non Klapalek 1905) (subjectve synonyms: japonica [A.] = aino [P.])

Nominal taxa included: 

Neobaetiella/g [g:1985]

Gratia/g [g:1992]


References. Novikova & Kluge 1987: * *;


Autapomorphies of Baetiella/g1.

(1) 2nd segment of labial palp lacks muscle, elongate, without apical projection; at the same time boundary between second and 3rd segments retained (Kluge 1997: Tab.11:12) (see Index of characters [1.1.58]).

(2) Larva is adapted to inhabitancy in rapid waters, its body Acentrella-like, i.e. with flattened ventral and convex dorsal side, thorax relatively large in comparison with abdomen (non-unique character – see Index of characters [1a]); paracercus is vestigial, 1-segmented (non-unique character – see Index of characters [1.3.64]); cerci are long, swimming setae are vestigial or lost (but at least vestigial bases of swimming setae are retained at least on middle segments). Dorsal side of tibia with a stripe of densely and irregularly situated long thin setae (Kluge 1997: Tab.11:13) (unlike Acentrella/g1, these setae do not form a regular straight row).

() Imago and subimago in rest keep their wings spread by sides (but being disturbed turn their wings up). (The character is known for two species only – tuberculatum [P.] and muchei [P.].) Majority of other Turbanoculata in rest keep their wings turning up, only selected species of Procloeon/g2 keep their wings in V-like position or spread by sides. Besides this, spread position of wings is characteristic for Ephemeroptera with modified pterothorax – for Caenoptera and for short-living Ephemeroptera with vestigial legs.

Characters of Baetiella/g1 of unclear phylogenetic status. 

() Larval abdominal terga lack wide scales [see Turbanoculata (12)] (see Index of characters [1.3.4]).

() Hind wings are lost [see Turbanoculata (10)]. Non-unique apomorphy (see Index of characters [2.2.59]).

Plesiomorphies of Baetiella/g1.

Unlike Acentrella/g1, ... imaginal and subimaginal anteronotal protuberance of mesonotum is low.

Variable characters of Baetiella/g1.  

Larval abdominal terga can be without projections (japonica [B.]), or with unpaired median projections near posterior margin (e.g., tuberculatum [P.]), or with paired submedian projections near posterior margin (bispinosa [P.], macani [N.]); the same in some other mayflies (see Index of chasracters [1.3.3]). 

Usually all tergalii I–VII are present, but in macani [N.] tergalii VII are lost; in the same species, a gill process is present medially to base of each coxa (the same in some other Turbanoculata – see Index of characters [1.2.12]).

Size. Fore wing length 4–7 mm(see Tetramerotarsata).

Distribution. Eastern and Middle Asia (to the west up to Tien-Shan).


Nominal species in Baetiella/g1:

aino Marsumura 1931 [Pseudocloeon] — typus nominis Matsumuracloeon — syn.subj. japonica [Acentrella]

armata Braasch 1983 [Baetiella]  

ausobskyi Braasch 1983 [Baetiella]  

bispinosus Gose 1980 [Pseudocloeon]

imanishii Braasch 1983 [Baetiella]  

japonica Imanishi 1931 [Acentrella] — typus nominis Baetiella  

lanpingensis Shi & Tong 2015 [Baetiella]

macani Müller-Liebenau 1985 [Neobaetiella]  

marginata Braasch 1983 [Baetiella]  

muchei Braasch 1978 [Pseudocloeon]  --/

narumonae Boonsong & Thomas & Sangpradub 2002 [Gratia]   --/

nosagawaensis Gose 1965 [Baetiella] — syn.subj. tuberculatum [Pseudocloeon]

sexta Shi & Tong 2015 [Baetiella]

sororculaenadinae Thomas 1992 [Gratia]  

spathae Shi & Tong 2015 [Baetiella]

trispinata Tong & Dudgeon 2000 [Baetiella]  

tuberculatum Kazlauskas 1963 [Pseudocloeon (Baetiella)]  --/

uenoi Müller-Liebenau 1985 [Neobaetiella] — typus nominis Neobaetiella — syn.subj. tuberculatum [Pseudocloeon]


Examined also:


See also:

Baetovectata INCERTAE SEDIS

Turbanoculata INCERTAE SEDIS